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The staggered air pressure just further ensures that the indexing pattern is in effect. In other words if a tire is the largest of the bunch @5 psi, then it will definitely be the largest at a little higher psi. Not to mention that slightly larger tires in the center help to make turning easier (same theory as the avenger's banana frame).
Indexing the tires ensures that the chains do not try to force any of the tensioners in the opposite direction than they are trying to work (when you drive your machine in the forward direction). That's why indexing should happen if you're running any track style. What happens next is that the center tires (even after indexing) then have to overdrive the track (different track styles to different degrees) to scrub-out their imperfect-from-the-factory slightly larger size. This is the part that wastes power and increases drivetrain load and is the part can be eliminated by track tuners. Youll see power gain and reduced load with tuners but they are not mandatory...so long as you index your tires. Fully indexing tires is mandatory if you don't use them, but tuners give you more flexibility regarding tire placement. You will be able to substitute "larger pairs" with "smaller pairs" when swapping corner tires and center tires and won't have to worry about the track prematurely rotating any of the tires, which is what improperly loads the chain tensioners. Tuners let you adjust center air pressure for ride and also allow you to effectively loosen or tighten your track (because the corner tires exclusively determine track length) by swapping tire positions without worrying about damaging tensioners. Sometimes the difference in tire circumfrence can effectively change your required track length by 1-2" simply based on the size of tires that are placed on the corner axles. The power gain (reduced load w/o slipping requirement) with tuners is most noticeable above 5 mph but is extremely beneficial at all speeds.
So, to repeat, if you're not going to use tuners, then definitely make sure to at least index tires per argo specs.
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